Methods:
DRAGNet will consist of 5 completely randomized blocks each with 5, 5x5m plots with each receiving one of the following treatments: long-term nutrient addition (10+ years), physical disturbance (3 sequential years of removal of aboveground vegetation, litter, rhizomes and tilling of the soil), nutrient addition combined with physical disturbance; short-term nutrient addition (5 years); or no treatments (controls).